doi: 10.24272/j.issn.2095-8137.2022.441

• Cheng-Liang Wang¹ , 
• Liang Chen⁶ , 
• Gang He² , 
• Xiao-Dong Yan⁷ , 
• Tong Wu¹ , 
• Wei Li² , 
• Hai-Tao Zhao¹ , 
• Yi Ren¹ , 
• Wei-Wei Fu¹ , 
• Yan Wang¹ , 
• He Zhang² , 
• Xiao-Wei Wang¹ , 
• Wei-Feng Wang³ , 
• Hong-Tao Zhao⁶ , 
• Ru-Liang Pan^{2, 4, 5 ,  ,}  , 
• Bao-Guo Li^{2 ,  ,} 

• 1.

  Shaanxi Key Laboratory of Animal Conservation, Shaanxi Institute of Zoology, Xi’an, Shaanxi 710032, China

• 2.

  Shaanxi Key Laboratory of Animal Conservation, College of Life Sciences, Northwest University, Xi’an, Shaanxi 710069, China

• 3.

  Shaanxi Nature Reserve and Wildlife Management Station, Shaanxi Forestry Bureau, Xi’an, Shaanxi 710082, China

• 4.

  School of Anatomy, Physiology and Human Biology, University of Western Australia, Western Australia 6907, Australia

• 5.

  International Centre of Biodiversity and Primate Conservation, Dali University, Dali, Yunnan 671003, China

• 6.

  Shaanxi Niubeiliang National Nature Reserve, Xi’an, Shaanxi 710100, China

• 7.

  Management Bureau of Shaanxi Changqing National Nature Reserve, Hanzhong, Zhejiang 723000, China

Permission for field survey in the Qinling Mountains was granted by the Shaanxi Forestry Bureau.
Supplementary data to this article can be found online.
The authors declare that they have no competing interests.
The idea for this study originally came from B.G.L., R.L.P., and C.L.W. Field data were collected by G.H., L.C., X.D.Y, T.W., W.L, H.T.Z., and H.Z. Historical distribution data were collected by Y.R., W.W.F., Y.W., and X.W.W. Statistical analyses were performed by C.L.W. The manuscript was prepared by C.L.W. and W.F.W. with edits and additional suggestions for data analysis and interpretation provided by B.G.L. and R.L.P. The project and all guidance were coordinated by W.F.W. All authors read and approved the final version of the manuscript.
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