[1] Andrews E. 2017. 7 Influential African Empires. From ancient Sudan to medieval Zimbabwe, get the facts on seven African kingdoms that made their mark on history. A&E Television Networks, LLC. URL: https://www.history.com/.amp/news/7-influential-african-empires.
[2] Applied Biosystems. 2009. DNA Sequencing by Capillary Electrophoresis, Applied BiosystemsChemistry Guide. Thermo Fisher Scientific.
[3] Bandelt HJ, Forster P, Röhl A. 1999. Median-joining networks for inferring intraspecific phylogenies. Molecular Biology and Evolution, 16(1): 37−48. doi:  10.1093/oxfordjournals.molbev.a026036
[4] Beja-Pereira A, England PR, Ferrand N, Jordan S, Bakhiet AO, Abdalla MA, Mashkour M, Jordana J, Taberlet P, Luikart G. 2004. African origins of the domestic donkey. Science, 304(5678): 1781. doi:  10.1126/science.1096008
[5] Blench RM. 2000. A History of Donkeys, Wild Asses and Mules in Africa. London, UK: University College London Press.
[6] Cardoso, J L, Vilstrup JT, Eisenmann V, Orlando L. 2013. First evidence of Equus asinus L. in the Chalcolithic disputes the Phoenicians as the first to introduce donkeys into the Iberian Peninsula. Journal of Archaeological Science, 40(12): 4483−4490. doi:  10.1016/j.jas.2013.07.010
[7] Chen SY, Zhou F, Xiao H, Sha T, Wu SF, Zhang YP. 2006. Mitochondrial DNA diversity and population structure of four Chinese donkey breeds. Animal. Genetics, 37(4): 427−429. doi:  10.1111/j.1365-2052.2006.01486.x
[8] Chen X, Ni G, He K, Ding ZL, Li GM, Adeola AC, Murphy RW, Wang WZ, Zhang YP. 2016. An improved de novo pipeline for enrichment of high diversity mitochondrial genomes from Amphibia to high-throughput sequencing. BioRxiv: 080689.
[9] Clutton-Brock J. 1992. Horse Power: A History of the Horse and the Donkey in Human Societies. Cambridge, USA: Harvard University Press.
[10] Drummond AJ, Rambaut A. 2007. BEAST: Bayesian evolutionary analysis by sampling trees. BMC Evolutionary. Biology, 7: 214. doi:  10.1186/1471-2148-7-214
[11] Endicott P, Ho SYW, Metspalu M, Stringer C. 2009. Evaluating the mitochondrial timescale of human evolution. Trends in Ecology and Evolution, 24(9): 515−521. doi:  10.1016/j.tree.2009.04.006
[12] Epstein H. 1971. The Origin of the Domestic Animals of Africa. New York, USA: African Publishing Corporation (APC).
[13] Excoffier L, Lischer HEL. 2010. Arlequin suite ver3.5: a new series of programs to perform population genetics analyses under Linux and Windows. Molecular Ecology Resources, 10(3): 564−567. doi:  10.1111/j.1755-0998.2010.02847.x
[14] Ge QL, Lei CZ, Jiang YQ, Chen H, Zhang W, Dang RH, Zhen HL, Zhang AL, Li TP. 2007. Genetic diversity and origins of Chinese donkey revealed by mitochondrial D-loop sequencies. Acta Veterinaria et Zootechnica Sinica, 38(7): 641−645.
[15] Han L, Zhu S, Ning C, Cai D, Wang K, Chen Q, Hu S, Yang J, Shao J, Zhu H, Zhou H. 2014. Ancient DNA provides new insight into the maternal lineages and domestication of Chinese donkeys. BMC Evolutionary Biology, 14: 246. doi:  10.1186/s12862-014-0246-4
[16] Herrera RJ, Garcia-Bertrand R. 2018. The Bantu Expansion. Ancestral DNA, human origins, and migrations. Academic Press, 395–432.
[17] Hiernaux J. 1968. Bantu expansion: the evidence from physical anthropology confronted with linguistic and archaeological evidence. The Journal of African History, 9(4): 505−515. doi:  10.1017/S0021853700009014
[18] Jordana J, Ferrando A, Miró J, Goyache F, Loarca A, Martinez López OR, Canelón JL, Stemmer A, Aguirre L, Lara MAC, Álvarez LA, Llambí S, Gómez N, Gama LT, Nóvoa MF, Martínez RD, Pérez E, Sierra A, Contreras MA, Guastella AM, Marletta D, Arsenos G, Curik I, Landi V, Martinez A, Delgado JV. 2016. Genetic relationships among American donkey populations: insights into the process of colonization. Journal of Animal Breeding and Genetics, 133(2): 155−164. doi:  10.1111/jbg.12180
[19] Kefena E, Dessie T, Tegegne A, Beja-Pereira A, Yusuf Kurtu M, Rosenbom S, Han JL. 2014. Genetic diversity and matrilineal genetic signature of native Ethiopian donkeys (Equus asinus) inferred from mitochondrial DNA sequence polymorphism. Livestock Science, 167: 73−79. doi:  10.1016/j.livsci.2014.06.006
[20] Kimura B, Marshall FB, Chen S, Rosenbom S, Moehlman PD, Tuross N, Sabin RC, Peters J, Barich B, Yohannes H, Kebede F, Teclai R, Beja-Pereira A, Mulligan CJ. 2011. Ancient DNA from Nubian and Somali wild ass provides insights into donkey ancestry and domestication. Proceedings of the Royal Society B: Biological Sciences, 278(1702): 50−57. doi:  10.1098/rspb.2010.0708
[21] Lindsay EH, Opdyke ND, Johnson NM. 1980. Pliocene dispersal of the horse Equus and late Cenozoic mammalian dispersal events. Nature, 287: 135−138. doi:  10.1038/287135a0
[22] Ma XY. 2018. Global Perspective of Phylogeography and Demongraphic History of Equus asinus Based on Mitochondrial Analysis. Master thesis, Yunnan University, China.
[23] Mark JJ. 2011. Punt. Ancient History Encyclopedia. https://www.ancient.eu/punt/.
[24] Marshall F. 2000. The Origins and Development of African Livestock. London: University College London Press.
[25] Marshall F. 2007. Rethinking Agriculture: Archeological and Ethnoarcheological Perspectives. Walnut Creek, CA, USA: Left Coast Press.
[26] Meutchieye F, Kwalar NN, Nyock RAF. 2017. Donkey Husbandry and Production Systems. Current Donkey Production and Functionality · Relationship with Humans. F. J. N. González, UCO Press.
[27] Mukhtār MJa-D . 1981. Ancient Civilizations of Africa. London: Heinemann Educational Books.
[28] Pérez-Pardal L, Grizelj J, Traoré A, Cubric-Curik V, Arsenos G, Dovenski T, Marković B, Fernández I, Cuervo M, Álvarez I, Beja-Pereira A, Curik I, Goyache F. 2014. Lack of mitochondrial DNA structure in Balkan donkey is consistent with a quick spread of the species after domestication. Animal Genetics, 45(1): 144−147. doi:  10.1111/age.12086
[29] R-Core-Team. 2019. R: A Language and Environment for Statistical Computing. R Foundation for Statistical Computing. https://www.R-project.org/.
[30] Rogers AR, Harpending H. 1992. Population growth makes waves in the distribution of pairwise genetic differences. Molecular Biology and Evolution, 9(3): 552−569.
[31] Rosenbom S, Costa V, Al-Araimi N, Kefena E, Abdel-Moneim AS, Abdalla MA, Bakhiet A, Beja-Pereira A. 2015. Genetic diversity of donkey populations from the putative centers of domestication. Animal Genetics, 46(1): 30−36. doi:  10.1111/age.12256
[32] Rossel S, Marshall F, Peters J, Pilgram T, Adams MD, O'Connor D. 2008. Domestication of the donkey: timing, processes, and indicators. Proceedings of the National Academy of Sciences of the United States of America, 105(10): 3715−3720. doi:  10.1073/pnas.0709692105
[33] Rozas J, Ferrer-Mata A, Sánchez-DelBarrio JC, Guirao-Rico S, Librado P, Ramos-Onsins SE, Sánchez-Gracia A. 2017. DnaSP 6: DNA sequence polymorphism analysis of large data sets. Molecular Biology and Evolution, 34(12): 3299−3302. doi:  10.1093/molbev/msx248
[34] Shi NN, Fan L, Yao YG, Peng MS, Zhang YP. 2014. Mitochondrial genomes of domestic animals need scrutiny. Molecular Ecology, 23(22): 5393−5397. doi:  10.1111/mec.12955
[35] Smith DG, Pearson RA. 2005. A review of the factors affecting the survival of donkeys in semi-arid regions of Sub-Saharan Africa. Tropical Animal Health and Production, 37(S1): 1−19. doi:  10.1007/s11250-005-9002-5
[36] South A. 2011. Rworldmap: a new R package for mapping global data. The R Journal, 3(1): 35−43. doi:  10.32614/RJ-2011-006
[37] Starkey P. 2000. The History of Working Animals in Africa. London, UK: University College London Press.
[38] Sun Y, Jiang Q, Yang C, Wang X, Tian F, Wang Y, Ma Y, Ju Z, Huang J, Zho X u, Zhong J, Wang C. 2016. Characterization of complete mitochondrial genome of Dezhou donkey (Equus asinus) and evolutionary analysis. Current Genetics, 62(2): 383−390. doi:  10.1007/s00294-015-0531-9
[39] Tamura K, Stecher G, Peterson D, Filipski A, Kumar S. 2013. MEGA6: molecular evolutionary genetics analysis version6.0. Molecular Biology and Evology, 30(12): 2725−2729. doi:  10.1093/molbev/mst197
[40] Thorvaldsdóttir H, Robinson JT, Mesirov JP. 2013. Integrative Genomics Viewer (IGV): high-performance genomics data visualization and exploration. Briefings Bioinformatics, 14(2): 178−192. doi:  10.1093/bib/bbs017
[41] Xia X, Yu J, Zhao X, Yao Y, Zeng L, Ahmed Z, Shen S, Dang R, Lei C. 2019. Genetic diversity and maternal origin of Northeast African and South American donkey populations. Animal Genetics, 50(3): 266−270. doi:  10.1111/age.12774
[42] Xie CX. 1987. The Equus Breeds in China. Shanghai: Shanghai Science & Technical Publishers. (in Chinese)
[43] Xu X, Gullberg A, Arnason U. 1996. The complete mitochondrial DNA (mtDNA) of the donkey and mtDNA comparisons among four closely related mammalian species-pairs. Journal of Molecular Evolution, 43(5): 438−446. doi:  10.1007/BF02337515